These features by which letters are recognized have been revealed by Fiset et al. (
2008) using the Bubbles technique. Notably, they showed that these letter features coincide mostly with line terminations and, to a lesser extent, with horizontal lines (see also Chung, Tjan, & Lin,
2008; Lanthier, Risko, Stolz, & Besner,
2009; Szwed, Cohen, Qiao, & Dehaene,
2009). However, Fiset et al. (
2008) overlooked an important aspect of these letter features: They fall almost exclusively on ink regions of letter images (see Fiset et al.,
2008, figure 2; Fiset et al.,
2009). This, as well as the fact that it was overlooked, suggests that observers typically do not use letter features falling on paper regions of letter images in letter recognition. In other words, it appears that observers rely on ink features—regions of ink in letter images—but not on paper features—regions of paper (without ink) in letter images. For example, observers rely on the presence of an ink curve to recognize the letter ‘o'—an ink feature—presumably to distinguish it from the letter ‘c,' but do not use the lack of ascenders or descenders—paper features—to distinguish it from the letters ‘b,' ‘d,' ‘p,' ‘q,' and ‘g.'